Background
Invasion of reticulocytes by P. vivax parasites is mediated through interaction between the Duffy-binding protein (PvDBP) and Duffy antigen on reticulocytes. Region II within PvDBP (PvDBP_RII) is the domain that binds Duffy antigen, indicating its potential as a target for vaccination. A blood-stage P. vivax Controlled Human Malaria Infection (CHMI) model has been developed in Oxford (UK)1, enabling the testing of vaccine efficacy by CHMI.
Methods
Two formulations of vaccines targeting PvDBP_RII, a viral-vectored2 and a protein-in-adjuvant formulation3, were tested for safety, immunogenicity and efficacy in Phase I/IIa trials in healthy UK adults. The viral-vectored vaccines were administered at 0, 2 months (ChAd63, MVA) or 0, 17, 19 months (ChAd63, ChAd63, MVA). The protein vaccine PvDBP_RII in Matrix-M™ adjuvant (Novavax AB®) (DBP/M-M) was administered in a monthly regimen (0, 1, 2 months) or delayed dosing regimen (0, 1, 14 months). Delayed regimens were necessitated due to trial halts during the pandemic. Volunteers underwent CHMI at 1 month following their last vaccination in parallel with unvaccinated controls. Efficacy was assessed by comparison of the blood-stage parasite multiplication rate (PMR) during CHMI, with parasitaemia measured by qPCR.
Results
Both vaccine formulations were safe and immunogenic. The delayed dosing regimen of DBP/M-M elicited the strongest anti-PvDBP_RII-specific antibody response and following CHMI, demonstrated an average 47% reduction in PMR (95% CI 20-74%) compared to unvaccinated controls. Other vaccine regimens tested did not demonstrate a significant impact on parasite growth.
Conclusions
The DBP/M-M vaccine administered in a delayed dosing regimen demonstrated a significant reduction in parasite growth following blood-stage CHMI, and is the first P. vivax vaccine to show in vivo impact on parasitaemia in humans. The improvement in efficacy of this vaccine when given in a delayed rather than monthly dosing regimen highlights the importance of optimising timing in vaccination regimens.